Operation mode: Think of the operation modes as presets that matches normal use-cases. ' . 'Usually you want to stick with Varied image responses or perhaps CDN friendly. ' . 'The Tweaked mode has no presets. That is: Here, you can set all options.

' . '

Changing from ie. "Varied image responses" mode to "Tweaked" mode enables you to see the underlying options for that mode (and tweak them). ' . 'Changing back will override the tweaks (you will lose them).

' . '

You will never loose your converter configurations by changing mode

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In the "Varied image responses" mode, WebP Express creates redirection rules for images, such that a request for a jpeg will result in a webp – but only if the request comes from a webp-enabled browser. Note that not all CDN's handles varied responses well (see FAQ).

In "CDN friendly" mode, a jpeg is always served as a jpeg. Instead of varying the image response, WebP Express alters the HTML for webp usage.
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A couple of options are available for automatically triggering webp conversion.

The "No conversion" mode is for scenarios where you are using another plugin for converting images. Perhaps the other plugin doesn't have the redirection or alter HTML feature, or perhaps it doesn't do it as well as WebP Express does. PS: The two methods below works great in tandem.